『Foundations of Systematics and Biogeography』

David M. Williams and Malte C. Ebach

(2008年1月刊行,Springer-Verlag, New York, xviii+309 pp., ISBN:978-0-387-72728-8版元ページコンパニオン・ブログ

【目次】
Foreword (Gareth Nelson) vii
Prologue ix
Acknowledgments xi

1. Introduction - Systematics, Evolution and Classification 1

 1.1 The Evolution of “Cladistics” 2
  1.1.1 What Is Cladistics? 2
  1.1.2 What Is Cladistics - Again? 6
 1.2 Cladistic Analysis 9
  1.2.1 Cladograms and Trees 9
  1.2.2 Cladograms and Evidence 12
 1.3 Cladistic Classification 14
  1.3.1 Cladistic Classification and Phylogeny 14
  1.3.2 Cladistic Classification and the Perils of Paraphyly 17
 1.4 Overview 18

2. Systematics as Problem-Solving 21

 2.1 The Problem 21
 2.2 The Solurions 21
 2.3 Discovering Solutions 24

3. The Archetype 28

 3.1 The Dynamic Organism: Bildung 28
 3.2 The Beginning of Comparative Biology: Goethe's Archetype 29
 3.3 Visualizing the Archetype 30
 3.4 Metamorphosis 32
 3.5 Urhomologie 33
 3.6 Relating the Ideal Organism 34
 3.7 Transformation and Goethe's Archetype 35

4. Ernst Haeckel and Systematische Phylogenie 37

 4.1 Ernst Haeckel and Darwin 37
 4.2 Haeckel's Genealogical Oaks and Stick “Trees” 38
 4.3 Heinrich Georg Bronn: Trunks and Twigs 39
 4.4 Schleicher, Lingustics & Trees 45
 4.5 Haeckel and Palaeontological Truth 47

5. The German Development of Morphology: From Ernst Haeckel to Willi Hennig 53

 5.1 Post-Haeckelian Systematics 53
 5.2 After Haeckel: Spemann and Homology 55
 5.3 Adolf Naef and “Systematische Morphologie”(Systematic Morphology) 56
 5.4 Konrad Lorenz, Homology, and Systematics 63
 5.5 Willi Hennig and the Resurrection of Transformation Systematics 64
 5.6 A Postscript on Naef's Criteria: Patterson's Homology Testing 67
  5.6.1 The Similarity Test 69
  5.6.2 The Complement Relation 72
  5.6.3 Two Homologies 74
  5.6.4 The Conjunction Test 76
  5.6.5 The Congruence Test 78
  5.6.6 Rejection of Patterson's Tests 80

6. Pattern Cladistics 82

 6.1 Preamble 86
 6.2 Discovering Léon Croizat 87
 6.3 Erik Stensiö, Vertebrate Palaeontology, and the Birth of Cladistics 89
 6.4 Lars Brundin, Vertebrate Palaeontology, and the Growth of Cladistics 93
 6.5 Lars Brundin and Léon Croizat: Conflict over Origins 95
 6.6 The London Reaction: A Salmon, a Lungfish, a Cow, and the Vertebrate Palaeontologists 96
  6.6.1 Two Schools and Extinct Fishes 98
  6.6.2 A Salmon, a Lungfish, a Cow, and Some Vertebrate Palaeontologists 100
  6.6.3 Exhibitions and Cladistics 102
  6.6.4 Mammals and More “Dinosaur” Cladistics 103
  6.6.5 Creationism, Marxism, Gradualism 106
  6.6.6 Halstead and Evolution 107
  6.6.7 More Creationism 111
  6.6.8 Karl Popper and Cladistics 112
  6.6.9 Even More Creationists: “Remember, Remember the 5th of November, Gunpowder, Treason and Plot ...” 113
 6.7 The Beginnings in New York: Fossils and Reform 117
 6.8 New York Collaborations: Distributions and Historical Biogeography 119
 6.9 The Two Cladistics 122
 6.10 Overview 123

7. Homologues and Homology 126

 7.1 The Search of the Unit of Classification 126
 7.2 Homologues and Analogues 131
 7.3 Homology and Analogy 131
 7.4 Homology 133
 7.5 Homotype 134
 7.6 Analogy 135
 7.7 The Conceptual Split: Owen's Archetype and the Path to Transformation 136
 7.8 Parts and Their Meaning 137

8. Discovering Homologues 139

 8.1 Homologues and Criteria 139
 8.2 Homologues and Phenetics 141
 8.3 Homologues and Cladistics 146
 8.4 The Data Matrix 149
  8.4.1 The Matrix: A Short Historical Digression 152

9. Homology and Systematics 155

 9.1 Natural Systems, Affinity, and Analogy 155
 9.2 William Sharp MacLeay and “Affinity” and “Analogy” 155
 9.3 The Horae Entomologicae (MacLeay 1819-1821) and After 158
 9.4 Westwood's Affinity and Analogy 160
 9.5 Strickland's Affinity and Analogy 161
 9.6 Owen and Strickland on Homology and Analogy 162
 9.7 Homology and Relationship 163
 9.8 Homology, Analogy, Parts, and Wholes 165

10. Homology and Transformation 168

 10.1 The Process of Change 168
 10.2 Material Transformation 169
  10.2.1 Phylogenetic (Historical) Transformation 169
  10.2.2 Ontogenetic Transformation 171
  10.2.3 Material Transformation and the Conflict of Causes 173
 10.3 Logical Transformation 174
 10.4 Genealogical Causes 175
 10.5 Transformation Revisited 177

11. Character Conflict 184

 11.1 The Problem of Conflict 184
 11.2 Cladistic Analysis 187
  11.2.1 Parsimony 187
  11.2.2 Strength of Evidence 188
  11.2.3 Contradictory Evidence 188
  11.2.4 Strength of Evidence and Contradictory Evidence 190
  11.2.5 “Computerised” parsimony 191
  11.2.6 Partial Charcters 191
  11.2.7 Informative Symplesiomorphy 192
  11.2.8 Resolving Conflict 193
 11.3 The Data-Matrix — Again: Component Characters 194
  11.3.1 Compatibility 195
  11.3.2 Component Analysis (Nelson 1979, Nelson & Platnick 1981) 198
  11.3.3 General Component Analysis (Nelson 1979, Nelson & Platnick 1981: 305-323, Page 1989b: 177-180) 199
  11.3.4 Patterson's Congruent Characters (1980a, 1982a, 1988a, b) 201
  11.3.5 Consistent and Congruent Characters (Scotland 1992, 1997, Kitching et al. 1998) 203
  11.3.6 Comparison 203
  11.3.7 Component Solutions: Paralogy 204
  11.3.8 Components and Equivalents 204
  11.3.9 Components as Data 206
 11.4 Augumented Component Analysis 207
  11.4.1 The Meaning of Statements of Relationship 208

12. The Analyses of Relationships 210

 12.1 Three-Item Data 210
 12.2 The Relationship Between Three-Item Statements and Binary Characters 213
 12.3 The Relationship Between Three-Item Statements and Multi-State Characters 213
 12.4 Cladogram Length and Three-Item Statements 213
 12.5 Information Measures: How Good Is a Three-Item Cladogram? 213
 12.6 Minimal Trees 213
 12.7 Explanations 216
 12.8 “Reversals” 216
 12.9 Alternatives to Simulation 217
  12.9.1 “Nullius in Verba” (Nelson 1996): Outgroups, Polarity, and Data 220
  12.9.2 Precision and Simple Matrices (Platnick et al. 1996) 224
  12.9.3 Comparison of Cladograms and Single Non-Conflicting Characters 224
 12.10 Summary 227

13. Biogeographical Relationships, Evolution and Classification 228

 13.1 Prelude 228
 13.2 The Threefold Parallelism: Its Beginning 231
 13.3 Haeckel's Hypothetische Skizze des monophyletischen Ursprungs und der Verbreitung der 12 Menschen-Species von Lemurien aus über die Erde and the Concept of Chorology 233
 13.4 The Development of Chorology 236
 13.5 Vicariance Versus Dispersal: Another “False War” 240
 13.6 Origins 241
 13.7 Realms, Regions, and Provinces 244
 13.8 Agassiz's (1854) Geological Realms: The Natural Provinces of Mankind 246
 13.9 Regions, Homology, and Relationships 249
 13.10 Sclater, Huxley, and the Classification of Regions 249
 13.11 Croizat's Radical Realms: Ocean Basin and Cladograms 251
 13.12 The Threfold Parallelism: ... and Its End 253
 13.13 Systematic Biogeography: The Rediscovery of Classification 254
  13.13.1 Area Homology 255
  13.13.2 Defining the Biotic Area: Biotic Morphology or Taxonomy 255
  13.13.3 Towards Area Monophyly 255
  13.13.4 Area Cladistics: Interpreting Area Monophyly 257


Epilogue 259
References 262
Index 305